B. A. Nikolay, Review of West Nile and Usutu virus co-circulation in Europe: How much do transmission cycles overlap?, Trans. R. Soc. Trop. Med. Hyg, vol.109, pp.609-618, 2015.

S. David and A. Abraham, Epidemiological and clinical aspects on West Nile virus, a globally emerging pathogen, Infect. Dis, vol.48, pp.571-586, 2016.

M. L. Bunning, R. A. Bowen, B. Cropp, K. Sullivan, B. Davis et al., Experimental infection of horses with West Nile virus and their potential to infect mosquitoes and serve as amplifying hosts, Ann. N. Y. Acad. Sci, vol.951, pp.338-339, 2001.

B. Durand, A. L. Tran, G. Balança, and V. Chevalier, Geographic variations of the bird-borne structural risk of West-Nile virus circulation in Europe, PLoS ONE, vol.12, 2017.
URL : https://hal.archives-ouvertes.fr/hal-02625587

V. Chevalier, A. Dupressoir, A. L. Tran, C. Gottland, V. Grosbois et al., Environmental risk factors of West Nile infection in the Senegal River basin, Epidemiol. Infect, vol.138, pp.1601-1609, 2010.

L. Bian, L. Li, and G. Yan, Combining global and local estimates for spatial distribution of mosquito larval habitats, GIS Sci. Remote Sens, vol.43, pp.128-141, 2006.

G. Valiakos, K. Papaspyropoulos, A. Giannakopoulos, P. Birtsas, S. Tsiodras et al., Use of Wild Bird Surveillance, Human Case Data and GIS Spatial Analysis for Predicting Spatial Distributions of West Nile Virus in Greece, PLoS ONE, vol.9, 2014.

R. Bargaoui, S. Lecollinet, and R. Lancelot, Mapping the Serological Prevalence Rate of West Nile fever in Equids, Tunisia. Trans. Emerg. Dis, vol.62, pp.55-66, 2013.
URL : https://hal.archives-ouvertes.fr/hal-02641436

V. Rodriguez-prieto, B. Martinez-lopez, M. Martinez, M. J. Munoz, and J. M. Sanchez-vizcaino, Identification of suitable areas for West Nile virus outbreaks in equid populations for application in surveillance plans: The example of the Castile and Leon region of Spain, Epidemiol. Infect, vol.140, pp.1617-1631, 2012.

A. Sánchez-gómez, C. Amela, E. Fernández-carrión, M. Martínez-avilés, J. Sánchez-vizcaíno et al., Risk mapping of West Nile virus circulation in Spain, Acta Trop, vol.169, pp.163-169, 2015.

N. Johnson, M. De-marco, A. Giovannini, C. Ippoliti, M. Danzetta et al., Emerging Mosquito-Borne Threats and the Response from European and Eastern Mediterranean Countries, Int. J. Environ. Res. Public Health, vol.15, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02627058

I. García-bocanegra, J. Belkhiria, S. Napp, D. Cano-terriza, S. Jiménez-ruiz et al., Epidemiology and spatio-temporal analysis of West Nile virus in horses in Spain between, Transbound. Emerg. Dis, vol.65, pp.567-577, 2010.

C. Rizzo, C. Napoli, G. Venturi, S. Pupella, L. Lombardini et al., West Nile virus transmission: Results from the integrated surveillance system in Italy, Euro Surveill, vol.21, 2008.

A. Conte, L. Candeloro, C. Ippoliti, F. Monaco, F. De-massis et al., Spatio-Temporal Identification of Areas Suitable for West Nile Disease in the Mediterranean Basin and Central Europe, PLoS ONE, vol.10, 2015.

A. Fall, A. Diaïté, M. Seck, J. Bouyer, T. Lefrançois et al., West Nile virus transmission in sentinel chickens and potential mosquito vectors, Int. J. Environ. Res. Public Health, vol.10, pp.4718-4727, 2008.
URL : https://hal.archives-ouvertes.fr/hal-02646626

, Int. J. Environ. Res. Public Health, vol.17, 2020.

V. Chevalier, P. Reynaud, T. Lefrançois, B. Durand, F. Baillon et al., Predicting West Nile virus seroprevalence in wild birds in Senegal. Vector Borne Zoonotic Dis, vol.9, pp.589-596, 2009.
URL : https://hal.archives-ouvertes.fr/hal-02667755

J. Renaudet, C. Jan, J. Ridet, C. Adam, and Y. Robin, A serological survey of arboviruses in the human population of Senegal, Bull. Soc. Pathol. Exot. Filiales, vol.71, pp.131-140, 1978.

C. Mathiot, Y. Clerc, F. Rodhain, J. P. Digoutte, and P. Coulanges, West Nile virus in Madagascar, Arch Inst Pasteur Madagascar, vol.51, pp.113-123, 1984.

M. Maquart, S. Boyer, V. M. Rakotoharinome, J. Ravaomanana, M. L. Tantely et al., High Prevalence of West Nile Virus in Domestic Birds and Detection in 2 New Mosquito Species in Madagascar, PLoS ONE, vol.11, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01300687

B. Nikolay, M. Diallo, C. S. Boye, and A. A. Sall, Usutu virus in Africa. Vector Borne Zoonotic Dis, vol.11, pp.1417-1423, 2011.

C. Ochieng, J. Lutomiah, A. Makio, H. Koka, E. Chepkorir et al., Mosquito-borne arbovirus surveillance at selected sites in diverse ecological zones of Kenya, Virol. J, vol.10, 2007.

T. Ben-hassine, F. De-massis, P. Calistri, G. Savini, B. Belhaj-mohamed et al., First detection of co-circulation of West Nile and Usutu viruses in equids in the south-west of Tunisia, Transbound. Emerg. Dis, vol.61, pp.385-389, 2014.

M. Diagne, M. Ndione, N. Di-paola, G. Fall, A. Bedekelabou et al., Usutu Virus Isolated from Rodents in Senegal, vol.11, p.181, 2019.

H. Weissenbock, J. Kolodziejek, A. Url, and H. Lussy, Rebel-Bauder, B.; Nowotny, N. Emergence of Usutu virus, an African mosquito-borne flavivirus of the Japanese encephalitis virus group, Europe. Emerg Infect. Dis, vol.8, pp.652-656, 2002.

U. Ashraf, J. Ye, X. Ruan, S. Wan, B. Zhu et al., Usutu virus: An emerging flavivirus in Europe, vol.7, pp.219-238, 2015.

T. Bakonyi, K. Erdelyi, K. Ursu, E. Ferenczi, T. Csorgo et al., Emergence of Usutu virus in Hungary, J. Clin. Microbiol, vol.45, pp.3870-3874, 2007.

U. Ziegler, H. Jost, K. Muller, D. Fischer, M. Rinder et al., Epidemic Spread of Usutu Virus in Southwest Germany in 2011 to 2013 and Monitoring of Wild Birds for Usutu and West Nile Viruses. Vector Borne Zoonotic Dis, vol.15, pp.481-488, 2015.

S. Lecollinet, Y. Blanchard, C. Manson, S. Lowenski, E. Laloy et al., Dual Emergence of Usutu Virus in Common Blackbirds, Emerg. Infect. Dis, vol.22, 2015.

J. M. Rijks, M. L. Kik, R. Slaterus, R. Foppen, A. Stroo et al., Widespread Usutu virus outbreak in birds in the Netherlands, vol.21, p.30391, 2016.

G. Kemenesi, D. Buzas, B. Zana, K. Kurucz, B. Krtinic et al., First genetic characterization of Usutu virus from Culex pipiens mosquitoes Serbia, Infect. Genet. Evol, vol.63, pp.58-61, 2014.

A. Vazquez, M. Jimenez-clavero, L. Franco, O. Donoso-mantke, V. Sambri et al., Usutu virus: Potential risk of human disease in Europe, Euro Surveill, vol.16, 2011.

M. Garigliany, A. Linden, G. Gilliau, E. Levy, M. Sarlet et al., Infect. Genet. Evol, vol.48, pp.116-119, 2016.

B. Mannasse, E. Mendelson, L. Orshan, O. Mor, U. Shalom et al., Usutu Virus RNA in mosquitoes, Israel, Emerg. Infect. Dis, vol.23, pp.1699-1702, 2014.

N. Becker, H. Jost, U. Ziegler, M. Eiden, D. Hoper et al., Epizootic emergence of Usutu virus in wild and captive birds in Germany, PLoS ONE, vol.7, 2012.

M. Cle, S. Salinas, S. Lecollinet, C. Beck, S. Gutierrez et al., Usutu virus: A new threat?, Epidemiol. Infect, vol.147, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02414235

D. Engel, H. Jöst, M. Wink, J. Börstler, S. Bosch et al., Reconstruction of the Evolutionary History and Dispersal of Usutu Virus, a Neglected Emerging Arbovirus in Europe and Africa, vol.7, pp.1938-1953, 2016.

H. Weissenböck, T. Bakonyi, G. Rossi, P. Mani, and N. Nowotny, Usutu virus, Emerg. Infect. Dis, vol.19, pp.274-277, 1996.

B. Durand, G. Balança, T. Baldet, and V. Chevalier, A metapopulation model to simulate West Nile virus circulation in Western Africa, Southern Europe and the Mediterranean basin, Vet. Res, vol.32, p.41, 2010.
URL : https://hal.archives-ouvertes.fr/hal-00903145

, Mediterranean-Black Sea Migration Pathway, Birdlife International, p.16, 2018.

L. Zwarts, R. Bijlsma, J. Van-der-kamp, and E. Wymenga, Changing Sahel, p.564, 2009.

J. Cappelle, R. Servan-de-almeida, B. Fofana, M. Dakouo, G. Balança et al., Circulation of avian influenza viruses in wild birds in Inner Niger Delta, Mali. Influenza Other Respir, Viruses, vol.6, pp.240-244, 2012.
URL : https://hal.archives-ouvertes.fr/hal-02650413

J. Cappelle, O. Girard, B. Fofana, N. Gaidet, and M. Gilbert, Ecological modeling of the spatial distribution of wild waterbirds to identify the main areas where avian influenza viruses are circulating in the Inner Niger Delta, Ecohealth, vol.7, pp.283-293, 2010.

C. Bebay, Première Évaluation de la Structure et de L'importance Dusecteur Avicole Commercial et Familial en Afrique de l'Ouest; Food and Agriculture Organization, p.22, 2006.

L. Ferry, M. Mietton, L. Robison, and J. Erismann, Lac Aloatra à Madagascar-Passé, présent et futur. Zeitschrift Fur Geomorphologie, vol.53, pp.299-318, 2009.

M. Pidgeon, An Ecological Survey of Lake Alaotra and Selected Wetlands of Central and Eastern Madagascar in Analyzing the Demise of Madagascar Pochard Aythya innotata

. Wwf/missouri-botanical-garden, , 1996.

O. Langrand, Delachaud et Niestlé; Delachaud and Niestlé: Lausanne, Suisse, p.432, 1995.

H. Rasamoelina-andriamanivo, R. Lancelot, O. F. Maminiaina, T. F. Rakotondrafara, M. Jourdan et al., Risk factors for avian pests in smallholder farming systems, Madagascar highlands, Prev. Vet. Med, vol.104, pp.114-124, 2011.

S. Molia, K. Samaké, A. Diarra, M. Sidibé, L. Doumbia et al., Avian influenza and Newcastle disease in three risk areas for H5N1 highly pathogenic avian influenza in Mali, Avian Dis, vol.55, pp.650-658, 2007.

C. Beck, I. Leparc-goffart, D. Desoutter, E. Deberge, H. Bichet et al., Serological evidence of infection with dengue and Zika viruses in horses on French Pacific Islands, PLoS Negl. Trop. Dis, vol.13, 2019.
URL : https://hal.archives-ouvertes.fr/inserm-02313831

. Oie, Manual of Diagnostic Tests and Vaccines for Terrestrial Animals, 2018.

R. Development-core and . Team, R: A Language and Environment for Statistical Computing; R Foundation for Statistical Computing, 2015.

M. Tantely, S. Goodman, T. Rakotondranaivo, and S. Boyer, Review of West Nile virus circulation and outbreak risk in Madagascar: Entomological and ornithological perspectives, Parasite, vol.23, p.49, 2016.

E. Cardinale, C. Bernard, S. Lecollinet, V. Rakotoharinome, J. Ravaomanana et al., West Nile virus infection in horses, Comp. Immunol. Microbiol. Infect. Dis, vol.53, pp.45-49, 2017.
URL : https://hal.archives-ouvertes.fr/pasteur-01675153

J. Morvan, L. Chin, D. Fontenille, I. Rakotoarivony, and P. Coulanges, The prevalence of West Nile virus-antibodies among children (5-20 years old) in Madagascar, Bull. Soc. Pathol. Exot, vol.84, pp.225-234, 1991.

S. Kalaiyarasu, N. Mishra, R. Khetan, and V. Singh, Serological evidence of widespread West Nile virus and Japanese encephalitis virus infection in native domestic ducks (Anas platyrhynchos var domesticus) in Kuttanad region, Kerala. India. Comp. Immunol. Microbiol. Infect. Dis, vol.48, pp.61-68, 2016.

S. Chaintoutis, S. Gewehr, S. Mourelatos, and C. Dovas, Serological monitoring of backyard chickens in Central Macedonia-Greece can detect low transmission of West Nile virus in the absence of human neuroinvasive disease cases, Acta Trop, vol.163, pp.26-31, 2016.

D. Fontenille, Transmission cycles of arboviruses in Madagascar. Arch Inst Pasteur Madag, vol.55, pp.7-317, 1989.

J. Yeh, J. Park, and E. Ostlund, Serologic evidence of West Nile Virus in wild ducks captured in major inland resting sites for migratory waterfowl in South Korea, Vet. Microbiol, vol.154, pp.96-103, 2011.

M. Saito, Y. Osa, and M. Asakawa, Antibodies to flaviviruses in wild ducks captured in Hokkaido, Japan: Risk assessment of invasive flaviviruses. Vector Borne Zoonotic Dis, vol.9, pp.253-258, 2009.

H. Shirafuj, K. Kanehira, M. Kubo, T. Shibahara, and T. Kamio, Experimental West Nile Virus Infection in Aigamo Ducks, a Cross between Wild Ducks (Anas platyrhynchos) and Domestic Ducks (Anas platyrhynchos var. domesticus), Avian Dis, vol.53, pp.239-244, 2009.

N. Komar, S. A. Langevin, S. R. Hinten, N. Nemeth, E. Edwards et al., Experimental infection of North American birds with the New-York 1999 strain of West Nile virus, Emerg. Infect. Dis, vol.9, pp.311-322, 2003.

V. O. Ezenwa, M. Godsey, R. King, and S. C. Guptill, Avian diversity and West Nile virus: Testing associations between biodiversity and infectious disease risk, Proc. R. Soc. B, vol.273, pp.109-117, 2006.

M. Tantely, G. Le-goff, S. Boyer, and D. Fontenille, An Updated Checklist of Mosquito Species (Diptera: Culicidae) from Madagascar, Parasite, vol.20, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01942689

D. A. Zimmerman, D. A. Turner, and D. J. Pearson, Birds of Kenya and Northern Tanzania, 1996.

M. Ciccozzi, S. Peletto, E. Cella, M. Giovanetti, A. Lai et al., Epidemiological history and phylogeography of West Nile virus lineage 2, Infect. Genet. Evol, vol.17, pp.46-50, 2013.

S. Larrieu, E. Cardinale, P. Ocquidant, M. Roger, R. Lepec et al., A fatal neuroinvasive West Nile virus infection in a traveller returning from Madagascar: Clinical, epidemiological and veterinary investigations, Am. J. Trop. Med. Hyg, vol.89, pp.211-213, 2013.

D. Fontenille, M. Traore-lamizana, M. Diallo, J. Thonnon, J. P. Digoutte et al., New vectors of Rift Valley fever in West Africa, Emerg. Infect. Dis, vol.4, pp.289-293, 1998.

V. Chevalier, R. Lancelot, A. Diaité, B. Mondet, B. Sall et al., Serological assessment of West Nile fever virus activity in the pastoral system of Ferlo (Senegal), Ann. N. Y. Acad. Sci, pp.216-225, 2006.

O. Cabre, J. P. Durand, A. Prange, J. Gomez, L. Maurizi et al., West Nile virus infection: Serological investigation among horses in France and in Africa, Med. Trop, vol.65, pp.439-443, 2005.

D. Safronetz, M. Sacko, N. Sogoba, K. Rosenke, C. Martellaro et al., Mali. Emerg. Infect. Dis, vol.22, pp.340-342, 2016.

N. J. Shahhosseini, S. Friedrich, M. Moosa-kazemi, M. Sedaghat, E. Kayedi et al., Host-feeding patterns of Culex mosquitoes in Iran. Parasites Vectors, vol.11, p.669, 2018.

K. Ergunay, F. Gunay, K. Erisoz, K. Oter, S. Gargari et al., Serological, molecular and entomological surveillance demonstrates widespread circulation of West Nile virus in Turkey, PLoS Negl. Trop. Dis, vol.8, 2014.

J. Meece, T. Kronenwetter-koepel, M. Vandermause, and K. Reed, West Nile virus infection in commercial waterfowl operation, Wisconsin. Emerg. Infect. Dis, vol.12, pp.1451-1453, 2006.

A. Kipp, J. Lehman, R. Bowen, P. Fox, M. Stephens et al., West Nile virus quantification in faeces of experimentally infected American and fish crows, Am. J. Trop. Med. Hyg, vol.75, pp.688-690, 2006.

R. Mclean, S. Ubico, D. Docherty, W. Hansen, L. Sileo et al., West Nile virus transmission and ecology in birds, Ann. N. Y. Acad. Sci, vol.951, pp.54-57, 2001.

A. Jaeger, S. Lecollinet, C. Beck, M. Bastien, M. Le-corre et al., Serological evidence for the circulation of flaviviruses in seabird populations of the western Indian Ocean, Epidemiol. Infect, vol.144, pp.652-660, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01180368

, Int. J. Environ. Res. Public Health, vol.17, 2020.

E. Ndiaye, D. Diallo, G. Fall, Y. Ba, O. Faye et al., Arboviruses isolated from the Barkedji mosquito-based surveillance system, BMC Infect. Dis, vol.18, pp.2012-2013, 2018.

L. Barzon, Ongoing and emerging arbovirus threats in Europe, J. Clin. Virol, vol.107, pp.38-47, 2018.