C. Augspurger, C. Karwautz, M. Mußmann, H. Daims, and T. J. Battin, Drivers of bacterial colonization patterns in stream biofilms, FEMS Microbiol. Ecol, vol.72, pp.47-57, 2010.

K. P. Aßhauer, B. Wemheuer, R. Daniel, and P. Meinicke, Tax4Fun: predicting functional profiles from metagenomic 16S rRNA data, Bioinformatics, vol.31, pp.2882-2884, 2015.

T. J. Battin, L. A. Kaplan, J. D. Newbold, X. Cheng, and C. Hansen, Effects of current velocity on the nascent architecture of stream microbial biofilms, 2003.

, Appl. Environ. Microbiol, vol.69, pp.5443-5452

K. Besemer, I. Hödl, G. Singer, and T. J. Battin, Architectural differentiation reflects bacterial community structure in stream biofilms, ISME J, vol.3, pp.1318-1324, 2009.

K. Besemer, G. Singer, R. Limberger, A. Chlup, G. Hochedlinger et al., Biophysical controls on community succession in stream biofilms, Appl. Environ. Microbiol, vol.73, pp.4966-4974, 2007.

J. Briand, A. Barani, C. Garnier, K. Réhel, F. Urvois et al., Spatio-temporal variations of marine biofilm communities colonizing artificial substrata including antifouling coatings in contrasted french coastal environments, Microb. Ecol, vol.74, pp.585-598, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01599314

J. Briand, I. Djeridi, D. Jamet, S. Coupé, C. Bressy et al., Pioneer marine biofilms on artificial surfaces including antifouling coatings immersed in two contrasting French Mediterranean coast sites, Biofouling, vol.28, pp.453-463, 2012.
URL : https://hal.archives-ouvertes.fr/hal-01362664

J. Bruzaud, J. Tarrade, A. Coudreuse, A. Canette, J. Herry et al., Flagella but not type IV pili are involved in the initial adhesion of Pseudomonas aeruginosa PAO1 to hydrophobic or superhydrophobic surfaces, Colloids Surf. B Biointerfaces, vol.131, pp.59-66, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01151646

J. G. Caporaso, J. Kuczynski, J. Stombaugh, K. Bittinger, F. D. Bushman et al., QIIME allows analysis of high-throughput community sequencing data, Nat. Methods, vol.7, pp.335-336, 2010.

C. Chagnot, M. A. Zorgani, T. Astruc, and M. Desvaux, Proteinaceous determinants of surface colonization in bacteria: bacterial adhesion and biofilm formation from a protein secretion perspective, Front. Microbiol, vol.4, p.303, 2013.
URL : https://hal.archives-ouvertes.fr/hal-01204318

C. Coclet, C. Garnier, F. Delpy, D. Jamet, G. Durrieu et al., Trace metal contamination as a toxic and structuring factor impacting ultraphytoplankton communities in a multicontaminated Mediterranean coastal area, Prog. Oceanogr, vol.163, pp.196-213, 2018.

C. Coclet, C. Garnier, G. Durrieu, D. Omanovic, S. D'onofrio et al., Changes in bacterioplankton communities resulting from direct and indirect interactions with trace metal gradients in an urbanized marine coastal area, Front. Microbiol, vol.10, p.257, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02053109

J. W. Costerton, Z. Lewandowski, D. E. Caldwell, D. R. Korber, and H. M. Lappinscott, Microbial biofilms, Annu. Rev. Microbiol, vol.49, pp.711-745, 1995.

B. G. Crespo, T. Pommier, B. Fernández-gómez, and C. Pedrós-alió, Taxonomic composition of the particle-attached and free-living bacterial assemblages in the Northwest Mediterranean sea analyzed by pyrosequencing of the 16S rRNA, vol.2, pp.541-552, 2013.

H. Dang and C. R. Lovell, Bacterial primary colonization and early succession on surfaces in marine waters as determined by amplified rRNA gene restriction analysis and sequence analysis of 16S rRNA genes, Appl. Environ. Microbiol, vol.66, pp.467-475, 2000.

H. Dang and C. R. Lovell, Microbial surface colonization and biofilm development in marine environments. Microbiol, Mol. Biol. Rev, vol.80, pp.91-138, 2016.

C. C. De-carvalho, Marine biofilms: a successful microbial strategy with economic implications. Front, Mar. Sci, vol.5, p.126, 2018.

E. F. Delong, D. G. Franks, A. , and A. L. , Phylogenetic diversity of aggregate-attached vs. free-living marine bacterial assemblages, Limnol. Oceanogr, vol.38, pp.924-934, 1993.

A. Dhariwal, J. Chong, S. Habib, I. L. King, L. B. Agellon et al., MicrobiomeAnalyst: a web-based tool for comprehensive statistical, visual and meta-analysis of microbiome data, Nucleic Acids Res, vol.45, 2017.

I. Douterelo, M. Jackson, C. Solomon, and J. Boxall, Microbial analysis of in situ biofilm formation in drinking water distribution systems: implications for monitoring and control of drinking water quality, Appl. Microbiol. Biotechnol, vol.100, pp.3301-3311, 2016.

W. M. Dunne, Bacterial adhesion: seen any good biofilms lately?, Clin. Microbiol. Rev, vol.15, pp.155-166, 2002.

T. H. Duong, J. Briand, A. Margaillan, and C. Bressy, Polysiloxanebased block copolymers with marine bacterial anti-adhesion properties, ACS Appl. Mater. Interfaces, vol.7, pp.15578-15586, 2015.

H. Elifantz, G. Horn, M. Ayon, Y. Cohen, and D. Minz, Rhodobacteraceae are the key members of the microbial community of the initial biofilm formed in Eastern Mediterranean coastal seawater, FEMS Microbiol. Ecol, vol.85, pp.348-357, 2013.

K. M. Eriksson, K. Sanli, H. Nilsson, A. Eiler, N. Corcoll et al., Triclosan changes community composition and selects for specific bacterial taxa in marine periphyton biofilms in low nanomolar concentration, PeerJ preprint, vol.6, pp.27125-27126, 2018.

D. P. Faith and A. M. Baker, Phylogenetic diversity (PD) and biodiversity conservation: some bioinformatics challenges, Evol. Bioinforma. Online, vol.2, pp.121-128, 2007.

L. Favre, A. Ortalo-magné, C. Pichereaux, A. Gargaros, O. Burlet-schiltz et al., Metabolome and proteome changes between biofilm and planktonic phenotypes of the marine bacterium Pseudoalteromonas lipolytica TC8, Biofouling, vol.34, pp.132-148, 2018.
URL : https://hal.archives-ouvertes.fr/hal-01760089

M. Fischer, G. Friedrichs, and T. Lachnit, Fluorescence-based quasicontinuous and In situ monitoring of biofilm formation dynamics in natural marine environments, Appl. Environ. Microbiol, vol.80, pp.3721-3728, 2014.

P. E. Galand, O. Pereira, C. Hochart, J. C. Auguet, and D. Debroas, A strong link between marine microbial community composition and function challenges the idea of functional redundancy, ISME J, vol.12, pp.2470-2478, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02002181

G. Galli and E. Martinelli, Amphiphilic polymer platforms: surface engineering of films for marine antibiofouling, Macromol. Rapid Commun, vol.38, p.1600704, 2017.

N. Gallois, B. Alpha-bazin, P. Ortet, M. Barakat, L. Piette et al., Proteogenomic insights into uranium tolerance of a chernobyl's microbacterium bacterial isolate, J. Proteomics, vol.177, pp.148-157, 2018.

J. F. Ghiglione, G. Mevel, M. Pujo-pay, L. Mousseau, P. Lebaron et al., Diel and seasonal variations in abundance, activity, and community structure of particle-attached and free-living bacteria in nw mediterranean sea, Microb. Ecol, vol.54, pp.217-231, 2007.

L. Girard, S. Peuchet, P. Servais, A. Henry, N. Charni-ben-tabassi et al., spatiotemporal dynamics of total viable Vibrio spp, NW mediterranean coastal area. Microbes Environ, vol.32, pp.210-218, 2017.

H. Grossart, Ecological consequences of bacterioplankton lifestyles: changes in concepts are needed: ecological consequences of bacterioplankton lifestyles, Environ. Microbiol. Rep, vol.2, pp.706-714, 2010.

J. Gutman, M. Herzberg, and S. L. Walker, Biofouling of reverse osmosis membranes: positively contributing factors of Sphingomonas, Environ. Sci. Technol, vol.48, pp.13941-13950, 2014.

L. Hall-stoodley, J. W. Costerton, and P. Stoodley, Bacterial biofilms: from the natural environment to infectious diseases, Nat. Rev. Microbiol, vol.2, pp.95-108, 2004.

J. J. Harrison, H. Ceri, and R. J. Turner, Multimetal resistance and tolerance in microbial biofilms, Nat. Rev. Microbiol, vol.5, pp.928-938, 2007.

D. Herbert-guillou, Influence of the hydrodynamics on the biofilm formation by mass transport analysis, Bioelectrochemistry, vol.53, pp.119-125, 2001.

B. D. Hoyle and J. W. Costerton, Bacterial resistance to antibiotics: the role of biofilms, Prog. Drug Res. Fortschritte Arzneimittelforschung Progres Rech. Pharm, vol.37, pp.91-105, 1991.

P. R. Jones, M. T. Cottrell, D. L. Kirchman, and S. C. Dexter, Bacterial community structure of biofilms on artificial surfaces in an estuary, Microb. Ecol, vol.53, pp.153-162, 2007.

T. Kiorboe, H. Grossart, H. Ploug, and K. Tang, Mechanisms and rates of bacterial colonization of sinking aggregates, Appl. Environ. Microbiol, vol.68, pp.3996-4006, 2002.

K. K. Kwon, J. Woo, S. Yang, J. Kang, S. G. Kang et al., Altererythrobacter epoxidivorans gen. nov., sp. nov., an epoxide hydrolase-active, mesophilic marine bacterium isolated from cold-seep sediment, and reclassification of erythrobacter luteolus Yoon et al. 2005 as Altererythrobacter luteolus comb. nov, Int. J. Syst. Evol. Microbiol, vol.57, pp.2207-2211, 2007.

S. Lambert, M. Tragin, J. Lozano, J. Ghiglione, D. Vaulot et al., Rhythmicity of coastal marine picoeukaryotes, bacteria and archaea despite irregular environmental perturbations, ISME J, vol.13, pp.388-401, 2018.

L. Costaouëc, T. Cérantola, S. Ropartz, D. Ratiskol, J. Sinquin et al., Structural data on a bacterial exopolysaccharide produced by a deep-sea Alteromonas macleodii strain, Carbohydr. Polym, vol.90, pp.49-59, 2012.

J. Lee, J. Nam, Y. Kim, K. Lee, and D. Lee, Bacterial communities in the initial stage of marine biofilm formation on artificial surfaces, J. Microbiol, vol.46, pp.174-182, 2008.

O. O. Lee, H. C. Chung, J. Yang, Y. Wang, S. Dash et al., Molecular techniques revealed highly diverse microbial communities in natural marine biofilms on polystyrene dishes for invertebrate larval settlement, Microb. Ecol, vol.68, pp.81-93, 2014.

M. Lejars, A. Margaillan, and C. Bressy, Fouling release coatings: a nontoxic alternative to biocidal antifouling coatings, Chem. Rev, vol.112, pp.4347-4390, 2012.

D. Lobelle, C. , and M. , Early microbial biofilm formation on marine plastic debris, Mar. Pollut. Bull, vol.62, 2011.

S. Louca, M. F. Polz, F. Mazel, M. B. Albright, J. A. Huber et al., Function and functional redundancy in microbial systems, Nat. Ecol. Evol, vol.2, pp.936-943, 2018.

S. Marceaux, C. Martin, A. Margaillan, and C. Bressy, Effects of accelerated ageing conditions on the mechanism of chemically-active antifouling coatings, Prog. Org. Coat, vol.125, pp.257-265, 2018.
URL : https://hal.archives-ouvertes.fr/hal-01878390

V. Michael, O. Frank, P. Bartling, C. Scheuner, M. Göker et al., Biofilm plasmids with a rhamnose operon are widely distributed determinants of the 'swim-or-stick' lifestyle in Roseobacters, ISME J, vol.10, pp.2498-2513, 2016.

B. Misson, C. Garnier, B. Lauga, D. H. Dang, J. Ghiglione et al., Chemical multi-contamination drives benthic prokaryotic diversity in the anthropized toulon bay, Sci. Total Environ, vol.556, pp.319-329, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01826037

T. Muthukrishnan, R. M. Abed, S. Dobretsov, B. Kidd, and A. A. Finnie, Long-term microfouling on commercial biocidal fouling control coatings, Biofouling, vol.30, pp.1155-1164, 2014.

T. K. Nam, M. B. Timmons, C. D. Montemagno, and S. M. Tsukuda, Biofilm characteristics as affected by sand size and location in fluidized bed vessels, Aquac. Eng, vol.22, pp.213-224, 2000.

R. Niederdorfer, H. Peter, and T. J. Battin, Attached biofilms and suspended aggregates are distinct microbial lifestyles emanating from differing hydraulics, Nat. Microbiol, vol.1, p.16178, 2016.

K. A. Nolte, J. Koc, J. M. Barros, K. Hunsucker, M. P. Schultz et al., Dynamic field testing of coating chemistry candidates by a rotating disk system, Biofouling, vol.34, pp.398-409, 2018.

S. Oberbeckmann, B. Kreikemeyer, and M. Labrenz, Environmental factors support the formation of specific bacterial assemblages on microplastics, Front. Microbiol, vol.8, p.2709, 2018.

S. Oberbeckmann, A. M. Osborn, and M. B. Duhaime, Microbes on a bottle: substrate, season and geography influence community composition of microbes colonizing marine plastic debris, PLoS One, vol.11, 2016.

A. E. Parada, D. M. Needham, and J. A. Fuhrman, Every base matters: assessing small subunit rRNA primers for marine microbiomes with mock communities, time series and global field samples: primers for marine microbiome studies, Environ. Microbiol, vol.18, pp.1403-1414, 2016.

L. Philippot, S. G. Andersson, T. J. Battin, J. I. Prosser, J. P. Schimel et al., The ecological coherence of high bacterial taxonomic ranks, Nat. Rev. Microbiol, vol.8, pp.523-529, 2010.

T. Pollet, L. Berdjeb, C. Garnier, G. Durrieu, C. Le-poupon et al., Prokaryotic community successions and interactions in marine biofilms: the key role of Flavobacteriia, FEMS Microbiol. Ecol, vol.94, p.83, 2018.
URL : https://hal.archives-ouvertes.fr/hal-01825993

B. Purevdorj, J. W. Costerton, and P. Stoodley, Influence of hydrodynamics and cell signaling on the structure and behavior of Pseudomonas aeruginosa biofilms, Appl. Environ. Microbiol, vol.68, pp.4457-4464, 2002.

A. Rieck, D. P. Herlemann, K. Jürgens, and H. Grossart, ParticleAssociated differ from free-living bacteria in surface waters of the baltic sea, Front. Microbiol, vol.6, p.1297, 2015.

A. Rochex, J. Godon, N. Bernet, and R. Escudié, Role of shear stress on composition, diversity and dynamics of biofilm bacterial communities, Water Res, vol.42, pp.4915-4922, 2008.

E. Romero, F. Peters, L. Arin, and J. Guillén, Decreased seasonality and high variability of coastal plankton dynamics in an urban location of the NW Mediterranean, J. Sea Res, vol.88, pp.130-143, 2014.

M. Salta, J. A. Wharton, Y. Blache, K. R. Stokes, and J. Briand, Marine biofilms on artificial surfaces: structure and dynamics, Environ. Microbiol, vol.15, pp.2879-2893, 2013.
URL : https://hal.archives-ouvertes.fr/hal-01364131

R. Schmieder and R. Edwards, Quality control and preprocessing of metagenomic datasets, Bioinformatics, vol.27, pp.863-864, 2011.

N. Segata, J. Izard, L. Waldron, D. Gevers, L. Miropolsky et al., Metagenomic biomarker discovery and explanation, Genome Biol, vol.12, p.60, 2011.

C. Sousa, P. Teixeira, and R. Oliveira, Influence of surface properties on the adhesion of staphylococcus epidermidis to acrylic and silicone, Int. J. Biomater, 2009.

P. Stoodley, I. Dodds, J. D. Boyle, and H. M. Scott, Influence of hydrodynamics and nutrients on biofilm structure, J. Appl. Microbiol, vol.85, pp.19-28, 1998.

P. Thomen, J. Robert, A. Monmeyran, A. Bitbol, C. Douarche et al., Bacterial biofilm under flow: first a physical struggle to stay, then a matter of breathing, PLoS One, vol.12, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01509809

Y. Tsai, Impact of flow velocity on the dynamic behaviour of biofilm bacteria, Biofouling, vol.21, pp.267-277, 2005.

N. A. Tujula, G. R. Crocetti, C. Burke, T. Thomas, C. Holmström et al., Variability and abundance of the epiphytic bacterial community associated with a green marine Ulvacean alga, vol.4, pp.301-311, 2010.

R. Vilchez, C. Pozo, M. A. Gomez, B. Rodelas, and J. Gonzalez-lopez, Dominance of sphingomonads in a copper-exposed biofilm community for groundwater treatment, Microbiology, vol.153, pp.325-337, 2007.

N. S. Webster and A. P. Negri, Site-specific variation in antarctic marine biofilms established on artificial surfaces, Environ. Microbiol, vol.8, 2006.

Y. Xia, Y. Kong, T. R. Thomsen, H. Nielsen, and P. , Identification and ecophysiological characterization of epiphytic proteinhydrolyzing saprospiraceae, Appl. Environ. Microbiol, vol.74, pp.2229-2238, 2008.

X. Xue, K. Zhang, F. Cai, J. Dai, Y. Wang et al., , 2012.

S. N. Altererythrobacter-xinjiangensis, isolated from desert sand, and emended description of the genus Altererythrobacter, Int. J. Syst. Evol. Microbiol, vol.62, pp.28-32

J. Yang, Y. Li, X. Guo, X. Liang, Y. Xu et al., The effect of carbon nanotubes and titanium dioxide incorporated in PDMS on biofilm community composition and subsequent mussel plantigrade settlement, Biofouling, vol.32, pp.763-777, 2016.

K. A. Zargiel and G. W. Swain, Static vs dynamic settlement and adhesion of diatoms to ship hull coatings, Biofouling, vol.30, pp.115-129, 2014.